Burke: Erythranthe primuloides vs. Erythranthe grandis

	Erythranthe primuloides	Erythranthe grandis
	primrose monkey-flower	large monkey-flower
Habit	Mat-forming perennial from long, thin rhizomes, the stem lax, up to 6 cm. long.	Rhizomatous perennials, occasionally rooting at nodes nearest base; stems usually 50-120 cm, erect or decumbent basally, branched, commonly hollow, densely hirsute to soft-hirtellous to long-hairy, hairs often crinkly, glandular or not, or sometimes villous-glandular without hirtellous hairs.
Leaves	Leaves opposite, crowded near the ground, short-hairy abaxially (underside), viscid to glabrous adaxially (upperside); leaves oblanceolate, sessile, nearly entire, 3-nerved from the base, 7-25 mm. long and 3-11 mm. wide.	Leaves basal and cauline, basal leaves generally not persistent; petioles 10-80 mm, becoming reduced distally; blade ovate to broadly elliptic, 25-60 mm long and 20-40 cm broad, palmate or nearly pinnate venation with 5-7 veins, base truncate to nearly cruneate to nearly cordate, margins crenulate to toothed, occasionally sublyrate near base, apex rounded to obtuse, surfaces of distalmost leaves hairy as stems.
Flowers	Flowers solitary on slender pedicels up to 10 cm. long arising from the leaf clusters; calyx narrow, 4-8 mm. long, mostly glabrous, the 5 teeth short, equal; corolla yellow, often dotted with maroon, 1-2 cm. long, scarcely bilabiate, the 5 lobes spreading, shallowly notched, the throat somewhat flaring; stamens 4.	Inflorescence racemose, bracteate, flowers 8-26; fruiting pedicels 10-35 mm, hairy as stems; calyx straight-erect or nodding at 45-100 degrees, ovate to bell-shaped, inflated, compressed along sagittal plane, 15-22 mm, hairy as stems, throat closing; corollas yellow with red spots inside, symmetric bilaterally, bilabiate; tube-throat widely funnel-shaped, 16-24 mm, protruding 10-15 mm beyond calyx margin, limb widely expanded; styles hirtellous; anthers not protruding, glabrous.
Fruit(s)	Capsule.	Capsules 8-12 mm, included.

	Erythranthe primuloides	Erythranthe grandis
Flowering time	June-August	May-August
Habitat	Wet meadows and boggy areas at middle to high elevations in the mountains.	Coastal areas including bluffs, dunes, wet cliff faces, marshes, ditches, lake and pond margins, and stream banks.
Distribution	Occurring on both sides of the Cascades crest in Washington; British Columbia to California, east to Montana, Utah, Arizona, and New Mexico. [WildflowerSearch map]	Occurring west of the Cascades crest in Washington; Washington to California. [WildflowerSearch map]
Origin	Native	Native
Conservation status	Not of concern	Not of concern
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis
Web links	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe grandis

primrose monkey-flower

large monkey-flower

Habit

Mat-forming perennial from long, thin rhizomes, the stem lax, up to 6 cm. long.

Rhizomatous perennials, occasionally rooting at nodes nearest base; stems usually 50-120 cm, erect or decumbent basally, branched, commonly hollow, densely hirsute to soft-hirtellous to long-hairy, hairs often crinkly, glandular or not, or sometimes villous-glandular without hirtellous hairs.

Leaves

Leaves opposite, crowded near the ground, short-hairy abaxially (underside), viscid to glabrous adaxially (upperside);

leaves oblanceolate, sessile, nearly entire, 3-nerved from the base, 7-25 mm. long and 3-11 mm. wide.

Leaves basal and cauline, basal leaves generally not persistent;

petioles 10-80 mm, becoming reduced distally;

blade ovate to broadly elliptic, 25-60 mm long and 20-40 cm broad, palmate or nearly pinnate venation with 5-7 veins, base truncate to nearly cruneate to nearly cordate, margins crenulate to toothed, occasionally sublyrate near base, apex rounded to obtuse, surfaces of distalmost leaves hairy as stems.

Flowers

Flowers solitary on slender pedicels up to 10 cm. long arising from the leaf clusters;

calyx narrow, 4-8 mm. long, mostly glabrous, the 5 teeth short, equal;

corolla yellow, often dotted with maroon, 1-2 cm. long, scarcely bilabiate, the 5 lobes spreading, shallowly notched, the throat somewhat flaring;

stamens 4.

Inflorescence racemose, bracteate, flowers 8-26; fruiting pedicels 10-35 mm, hairy as stems;

calyx straight-erect or nodding at 45-100 degrees, ovate to bell-shaped, inflated, compressed along sagittal plane, 15-22 mm, hairy as stems, throat closing;

corollas yellow with red spots inside, symmetric bilaterally, bilabiate;

tube-throat widely funnel-shaped, 16-24 mm, protruding 10-15 mm beyond calyx margin, limb widely expanded;

styles hirtellous;

anthers not protruding, glabrous.

Fruit(s)

Capsule.

Capsules 8-12 mm, included.