Burke: Erythranthe patula vs. Erythranthe caespitosa

	Erythranthe patula	Erythranthe caespitosa
	stalk-leaved monkey-flower	large mountain monkey-flower
Habit	Annuals with fibrous roots or a filiform-taproot; stems usually 5-15 cm, erect to ascending, straight or sharply bent at nodes, usually unbranched, covered with stalked glands, gland-tipped hairs 0.2-0.5 mm.	Rhizomatous perennials, rooting at nodes nearest the base, occasionally producing runners with creeping form and small leaves, forming colonies of mats, rhizomes threadlike; stems 3-10 cm, usually prostrate or with some curving upward distally, usually massed, round or somewhat flat, branching, may be glabrous or slightly hirtellous or with stalked glands.
Leaves	Leaves cauline, basal ones not persistent; petioles 8-25 mm; blade deltate or somewhat ovate to lanceolate, approximately 4-12 mm long and 3-10 mm broad, palmate venation with 3 veins, base rounded to cuneate-truncate, margins usually finely toothed, apex acute to obtuse, surfaces glabrous as stems.	Leaves both basal and cauline; petiolate, proximal petioles 2-5 mm, becoming sessile distally; blade commonly purplish beneath, orbicular to ovate to narrowly elliptic, leaves nearest base somewhat lyrate, 3-12 mm and becoming larger farther from base, palmate venation with 3 veins, base wedge-shaped, margins entire to barely finely toothed, apex obtuse, somewhat puberulent above and beneath, hairs minute and stipitate-glandular.
Flowers	Axillary flowers 1-10, emerging from nodes throughout; fruiting pedicels 10-25 mm, glandular as stems; calyx tubular, barely or not inflated, 5-6 mm, margins with distinct teeth or lobes, slightly stipitate-glandular to sparsely hirtellous, lobes pronounced, erect; corollas yellow, lower limb commonly with some red or brownish dots, symmetric radially or bilaterally, regular or weakly bilabiate; tube-throat funnel-shaped, 7-8 mm, protruding beyond calyx margin; lobes oblong, apex rounded to truncate; styles glabrous; anthers not protruding, glabrous.	Axillary flowers 1-3, from nodes farthest from base, usually solitary; fruiting pedicels generally 10-30 mm, somewhat villous with short gland-tipped hairs, occasionally hirtellous; calyx broadly bell-shaped, inflated, compressed along sagittal plane, 7-15 mm, glabrous or hirtellous or stipitate-glandular, throat closing, lobe pair nearest base curving upwards, other lobes 3-5 mm, prominently protruding; corollas yellow with dark red spots, symmetric bilaterally, bilabiate; tube-throat widely funnel-shaped to nearly cylindric, 15-18 mm, protruding beyond calyx margin; lower limb deflexed and spreading, upper limb curving upward, palate partially closed; styles slightly hirtellous; anthers not protruding, glabrous.
Fruits	Capsules 4-6 mm, included.	Capsules 4-5 mm, included.

	Erythranthe patula	Erythranthe caespitosa
Flowering time	May-July	July-September
Habitat	Vernally moist areas, seeps, and stream banks from the lowlands to the middle elevations.	Wet meadows and wet, rocky slopes at high elevations in the mountains.
Distribution	Occurring east of the Cascades crest in southeastern Washington; southeast Washington to adjacent northeast Oregon and adjacent west-central Idaho. [WildflowerSearch map]	Occurring in the Olympics and Cascades mountains of Washington; British Columbia south to Oregon, east to Idaho. [WildflowerSearch map]
Origin	Native	Native
Conservation status	Threatened in Washington (WANHP)	Not of concern
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis
Web links	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe caespitosa

stalk-leaved monkey-flower

large mountain monkey-flower

Habit

Annuals with fibrous roots or a filiform-taproot; stems usually 5-15 cm, erect to ascending, straight or sharply bent at nodes, usually unbranched, covered with stalked glands, gland-tipped hairs 0.2-0.5 mm.

Rhizomatous perennials, rooting at nodes nearest the base, occasionally producing runners with creeping form and small leaves, forming colonies of mats, rhizomes threadlike; stems 3-10 cm, usually prostrate or with some curving upward distally, usually massed, round or somewhat flat, branching, may be glabrous or slightly hirtellous or with stalked glands.

Leaves

Leaves cauline, basal ones not persistent;

petioles 8-25 mm;

blade deltate or somewhat ovate to lanceolate, approximately 4-12 mm long and 3-10 mm broad, palmate venation with 3 veins, base rounded to cuneate-truncate, margins usually finely toothed, apex acute to obtuse, surfaces glabrous as stems.

Leaves both basal and cauline; petiolate, proximal petioles 2-5 mm, becoming sessile distally;

blade commonly purplish beneath, orbicular to ovate to narrowly elliptic, leaves nearest base somewhat lyrate, 3-12 mm and becoming larger farther from base, palmate venation with 3 veins, base wedge-shaped, margins entire to barely finely toothed, apex obtuse, somewhat puberulent above and beneath, hairs minute and stipitate-glandular.

Flowers

Axillary flowers 1-10, emerging from nodes throughout; fruiting pedicels 10-25 mm, glandular as stems;

calyx tubular, barely or not inflated, 5-6 mm, margins with distinct teeth or lobes, slightly stipitate-glandular to sparsely hirtellous, lobes pronounced, erect;

corollas yellow, lower limb commonly with some red or brownish dots, symmetric radially or bilaterally, regular or weakly bilabiate;

tube-throat funnel-shaped, 7-8 mm, protruding beyond calyx margin;

lobes oblong, apex rounded to truncate;

styles glabrous;

anthers not protruding, glabrous.

Axillary flowers 1-3, from nodes farthest from base, usually solitary; fruiting pedicels generally 10-30 mm, somewhat villous with short gland-tipped hairs, occasionally hirtellous;

calyx broadly bell-shaped, inflated, compressed along sagittal plane, 7-15 mm, glabrous or hirtellous or stipitate-glandular, throat closing, lobe pair nearest base curving upwards, other lobes 3-5 mm, prominently protruding;

corollas yellow with dark red spots, symmetric bilaterally, bilabiate;

tube-throat widely funnel-shaped to nearly cylindric, 15-18 mm, protruding beyond calyx margin; lower limb deflexed and spreading, upper limb curving upward, palate partially closed;

styles slightly hirtellous;

anthers not protruding, glabrous.

Fruits

Capsules 4-6 mm, included.

Capsules 4-5 mm, included.