Burke: Erythranthe nasuta vs. Erythranthe pulsiferae

	Erythranthe nasuta	Erythranthe pulsiferae
	large-nose monkey-flower, snouted monkey-flower	candelabrum monkey-flower, Pulsifer's monkey-flower
Habit	Annuals with fibrous roots or a slender taproot; stems usually 2-35 cm, erect to ascending-erect or decumbent, unbranched or branched from nodes nearest base, 4-angled or occasionally weakly 4-winged, thin and wiry or hollow, glabrous except for small villous-glandular areas directly above nodes, occasionally with minute rigid hairs distally.	Annuals with shallow, fibrous roots; stems 5-12 cm, erect, not bent or curved at nodes, not branched or only slightly branched proximally, covered with minute stalked glands, gland-tipped hairs 0.1-0.3.
Leaves	Leaves basal and cauline or basal may not persist; petioles 3-35 mm from base to mid-stem with narrowly flanged bases, becoming sessile distally; blade reddish or purple-spotted beneath, somewhat ovate to nearly orbicular, approximately 10-49 mm long and 10-25 mm broad, leaves nearest base largest and persistent, palmate venation with 3-5 veins, base cuneate to truncate or nearly cordate, margins irregularly toothed or nearly lacerate, commonly doubly-toothed, 4-9 main teeth per side, sometimes sublacerate to sublyrate near base, apex acute to obtuse, surfaces glabrous or hirtellous with hairs that are dull, cylindrical, sharp-pointed, eglandular.	Leaves basal and cauline; petioles 2-9 mm with 3 distinct veins and 2 wings; blade elliptic-oblong to ovate or oblanceolate, 3-14 mm long and 2-9 mm broad, palmate venation with 3 veins, base cuneate to attenuate, margins finely toothed to entire, apex acute to obtuse, surfaces glandular-hairy as stems.
Flowers	Axillary flowers typically 2-12, emerging from nodes at mid- to ends of stems; fruiting pedicels usually 7-20 mm, longer than subtending leaves, becoming glabrous, occasionally glandular-villous on upper surface at axils; calyx nodding at 30-180 degrees, commonly purplish or purple-spotted, ovoid-campanulate, inflated, compressed across sagittal plane, usually 10-15 mm, hirtellous or appressed-hirtellous, or glabrous, slightly short-ciliate at sinuses, throat closing, upper lobe generally longer than lower lobe, slightly sickle shaped; corollas yellow, throat commonly with red spots, lower limb usually with a red splotch at base, symmetric bilaterally, weakly bilabiate; tube-throat widely cylindric, 8-12 mm, equal to or protruding 1-2 mm beyond calyx margin, limb expanded 6-12 mm; styles slightly rough or glabrous; anthers not protruding, glabrous.	Axillary flowers 1-9, emerging from mid-stem to distal nodes; fruiting pedicels horizontally spreading-curved, 12-38 mm, glandular-hairy as stems; calyx cylindric, somewhat inflated, 7-10 mm, margins toothed or lobed, glandular-hairy as stems, lobes pronounced, erect; corollas yellow, tube-throat, palate ridges, lower limb yellow to light yellow, lower limb with red dots or not, symmetric bilaterally, barely bilabiate; tube-throat funnel-shaped and 6-9 mm, protruding beyond calyx margin; lobes widely obovate to somewhat orbicular, apex rounded; styles glabrous; anthers not protruding, glabrous.
Fruits	Capsules 5-9 mm, included.	Capsules 5-8 mm, included.

	Erythranthe nasuta	Erythranthe pulsiferae
Flowering time	May-July	May-July
Habitat	Rocky or moist slopes, stream banks, and seeps.	Vernally moist areas, seeps, and stream banks from the lowlands to middle elevations.
Distribution	Occurring on both sides of the Cascades crest in Washington; southern British Columbia to Baja California, east to Idaho, Utah, and New Mexico. [WildflowerSearch map]	Occurring east of the Cascades crest in south-central Washington; south-central Washington to California, [WildflowerSearch map]
Origin	Native	Native
Conservation status	Not of concern	Sensitive in Washington (WANHP)
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. scouleri, E. suksdorfii, E. washingtonensis
Web links	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe pulsiferae

large-nose monkey-flower, snouted monkey-flower

candelabrum monkey-flower, Pulsifer's monkey-flower

Habit

Annuals with fibrous roots or a slender taproot; stems usually 2-35 cm, erect to ascending-erect or decumbent, unbranched or branched from nodes nearest base, 4-angled or occasionally weakly 4-winged, thin and wiry or hollow, glabrous except for small villous-glandular areas directly above nodes, occasionally with minute rigid hairs distally.

Annuals with shallow, fibrous roots; stems 5-12 cm, erect, not bent or curved at nodes, not branched or only slightly branched proximally, covered with minute stalked glands, gland-tipped hairs 0.1-0.3.

Leaves

Leaves basal and cauline or basal may not persist;

petioles 3-35 mm from base to mid-stem with narrowly flanged bases, becoming sessile distally;

blade reddish or purple-spotted beneath, somewhat ovate to nearly orbicular, approximately 10-49 mm long and 10-25 mm broad, leaves nearest base largest and persistent, palmate venation with 3-5 veins, base cuneate to truncate or nearly cordate, margins irregularly toothed or nearly lacerate, commonly doubly-toothed, 4-9 main teeth per side, sometimes sublacerate to sublyrate near base, apex acute to obtuse, surfaces glabrous or hirtellous with hairs that are dull, cylindrical, sharp-pointed, eglandular.

Leaves basal and cauline;

petioles 2-9 mm with 3 distinct veins and 2 wings;

blade elliptic-oblong to ovate or oblanceolate, 3-14 mm long and 2-9 mm broad, palmate venation with 3 veins, base cuneate to attenuate, margins finely toothed to entire, apex acute to obtuse, surfaces glandular-hairy as stems.

Flowers

Axillary flowers typically 2-12, emerging from nodes at mid- to ends of stems; fruiting pedicels usually 7-20 mm, longer than subtending leaves, becoming glabrous, occasionally glandular-villous on upper surface at axils;

calyx nodding at 30-180 degrees, commonly purplish or purple-spotted, ovoid-campanulate, inflated, compressed across sagittal plane, usually 10-15 mm, hirtellous or appressed-hirtellous, or glabrous, slightly short-ciliate at sinuses, throat closing, upper lobe generally longer than lower lobe, slightly sickle shaped;

corollas yellow, throat commonly with red spots, lower limb usually with a red splotch at base, symmetric bilaterally, weakly bilabiate;

tube-throat widely cylindric, 8-12 mm, equal to or protruding 1-2 mm beyond calyx margin, limb expanded 6-12 mm;

styles slightly rough or glabrous;

anthers not protruding, glabrous.

Axillary flowers 1-9, emerging from mid-stem to distal nodes; fruiting pedicels horizontally spreading-curved, 12-38 mm, glandular-hairy as stems;

calyx cylindric, somewhat inflated, 7-10 mm, margins toothed or lobed, glandular-hairy as stems, lobes pronounced, erect;

corollas yellow, tube-throat, palate ridges, lower limb yellow to light yellow, lower limb with red dots or not, symmetric bilaterally, barely bilabiate;

tube-throat funnel-shaped and 6-9 mm, protruding beyond calyx margin;

lobes widely obovate to somewhat orbicular, apex rounded;

styles glabrous;

anthers not protruding, glabrous.

Fruits

Capsules 5-9 mm, included.

Capsules 5-8 mm, included.