Burke: Erythranthe grandis vs. Erythranthe pulsiferae

	Erythranthe grandis	Erythranthe pulsiferae
	large monkey-flower	candelabrum monkey-flower, Pulsifer's monkey-flower
Habit	Rhizomatous perennials, occasionally rooting at nodes nearest base; stems usually 50-120 cm, erect or decumbent basally, branched, commonly hollow, densely hirsute to soft-hirtellous to long-hairy, hairs often crinkly, glandular or not, or sometimes villous-glandular without hirtellous hairs.	Annuals with shallow, fibrous roots; stems 5-12 cm, erect, not bent or curved at nodes, not branched or only slightly branched proximally, covered with minute stalked glands, gland-tipped hairs 0.1-0.3.
Leaves	Leaves basal and cauline, basal leaves generally not persistent; petioles 10-80 mm, becoming reduced distally; blade ovate to broadly elliptic, 25-60 mm long and 20-40 cm broad, palmate or nearly pinnate venation with 5-7 veins, base truncate to nearly cruneate to nearly cordate, margins crenulate to toothed, occasionally sublyrate near base, apex rounded to obtuse, surfaces of distalmost leaves hairy as stems.	Leaves basal and cauline; petioles 2-9 mm with 3 distinct veins and 2 wings; blade elliptic-oblong to ovate or oblanceolate, 3-14 mm long and 2-9 mm broad, palmate venation with 3 veins, base cuneate to attenuate, margins finely toothed to entire, apex acute to obtuse, surfaces glandular-hairy as stems.
Flowers	Inflorescence racemose, bracteate, flowers 8-26; fruiting pedicels 10-35 mm, hairy as stems; calyx straight-erect or nodding at 45-100 degrees, ovate to bell-shaped, inflated, compressed along sagittal plane, 15-22 mm, hairy as stems, throat closing; corollas yellow with red spots inside, symmetric bilaterally, bilabiate; tube-throat widely funnel-shaped, 16-24 mm, protruding 10-15 mm beyond calyx margin, limb widely expanded; styles hirtellous; anthers not protruding, glabrous.	Axillary flowers 1-9, emerging from mid-stem to distal nodes; fruiting pedicels horizontally spreading-curved, 12-38 mm, glandular-hairy as stems; calyx cylindric, somewhat inflated, 7-10 mm, margins toothed or lobed, glandular-hairy as stems, lobes pronounced, erect; corollas yellow, tube-throat, palate ridges, lower limb yellow to light yellow, lower limb with red dots or not, symmetric bilaterally, barely bilabiate; tube-throat funnel-shaped and 6-9 mm, protruding beyond calyx margin; lobes widely obovate to somewhat orbicular, apex rounded; styles glabrous; anthers not protruding, glabrous.
Fruits	Capsules 8-12 mm, included.	Capsules 5-8 mm, included.

	Erythranthe grandis	Erythranthe pulsiferae
Flowering time	May-August	May-July
Habitat	Coastal areas including bluffs, dunes, wet cliff faces, marshes, ditches, lake and pond margins, and stream banks.	Vernally moist areas, seeps, and stream banks from the lowlands to middle elevations.
Distribution	Occurring west of the Cascades crest in Washington; Washington to California. [WildflowerSearch map]	Occurring east of the Cascades crest in south-central Washington; south-central Washington to California, [WildflowerSearch map]
Origin	Native	Native
Conservation status	Not of concern	Sensitive in Washington (WANHP)
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. scouleri, E. suksdorfii, E. washingtonensis
Web links	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe pulsiferae

large monkey-flower

candelabrum monkey-flower, Pulsifer's monkey-flower

Habit

Rhizomatous perennials, occasionally rooting at nodes nearest base; stems usually 50-120 cm, erect or decumbent basally, branched, commonly hollow, densely hirsute to soft-hirtellous to long-hairy, hairs often crinkly, glandular or not, or sometimes villous-glandular without hirtellous hairs.

Annuals with shallow, fibrous roots; stems 5-12 cm, erect, not bent or curved at nodes, not branched or only slightly branched proximally, covered with minute stalked glands, gland-tipped hairs 0.1-0.3.

Leaves

Leaves basal and cauline, basal leaves generally not persistent;

petioles 10-80 mm, becoming reduced distally;

blade ovate to broadly elliptic, 25-60 mm long and 20-40 cm broad, palmate or nearly pinnate venation with 5-7 veins, base truncate to nearly cruneate to nearly cordate, margins crenulate to toothed, occasionally sublyrate near base, apex rounded to obtuse, surfaces of distalmost leaves hairy as stems.

Leaves basal and cauline;

petioles 2-9 mm with 3 distinct veins and 2 wings;

blade elliptic-oblong to ovate or oblanceolate, 3-14 mm long and 2-9 mm broad, palmate venation with 3 veins, base cuneate to attenuate, margins finely toothed to entire, apex acute to obtuse, surfaces glandular-hairy as stems.

Flowers

Inflorescence racemose, bracteate, flowers 8-26; fruiting pedicels 10-35 mm, hairy as stems;

calyx straight-erect or nodding at 45-100 degrees, ovate to bell-shaped, inflated, compressed along sagittal plane, 15-22 mm, hairy as stems, throat closing;

corollas yellow with red spots inside, symmetric bilaterally, bilabiate;

tube-throat widely funnel-shaped, 16-24 mm, protruding 10-15 mm beyond calyx margin, limb widely expanded;

styles hirtellous;

anthers not protruding, glabrous.

Axillary flowers 1-9, emerging from mid-stem to distal nodes; fruiting pedicels horizontally spreading-curved, 12-38 mm, glandular-hairy as stems;

calyx cylindric, somewhat inflated, 7-10 mm, margins toothed or lobed, glandular-hairy as stems, lobes pronounced, erect;

corollas yellow, tube-throat, palate ridges, lower limb yellow to light yellow, lower limb with red dots or not, symmetric bilaterally, barely bilabiate;

tube-throat funnel-shaped and 6-9 mm, protruding beyond calyx margin;

lobes widely obovate to somewhat orbicular, apex rounded;

styles glabrous;

anthers not protruding, glabrous.

Fruits

Capsules 8-12 mm, included.

Capsules 5-8 mm, included.