Burke: Arctostaphylos nevadensis vs. Arctostaphylos columbiana

	Arctostaphylos nevadensis	Arctostaphylos columbiana
	pinemat manzanita	bristly manzanita
Habit	Shrubs with spreading to decumbent stems often forming large mats or mounds, the branch tips to 2 dm. tall; bark brownish-red, stems puberulent, sometimes glandular.	Erect or spreading shrubs 1-3 m. tall, simple at the base, the old branches with purplish-red bark; young twigs grayish-pubescent, also with scattered bristles which may be glandular.
Leaves	Leaves oblong to spatulate, the tips acute or abruptly pointed; blades 1.5-3 cm. long, glabrous to puberulent on the margins and mid-rib, the base acute or narrowed abruptly to petioles 2-5 mm. long.	Leaves alternate, ovate or lanceolate to elliptic, the blades 2-5 cm. long and 1-2.5 cm. broad, pointed, finely grayish-puberulent, the petioles 5 cm. long.
Flowers	: Inflorescence a terminal, few-flowered raceme, each flower subtended by a bract about equal to the pedicel; flowers pink, about 5 mm. long; corolla urn-shaped, with 5 short lobes; stamens 10, the anthers opening by false terminal pores, each with 2 curved, reflexed, horn-like appendages; ovary superior, pubescent.	Inflorescence usually paniculate, densely pubescent, each flower subtended by a lanceolate bract 5-15 mm. long, exceeding the pedicels; flowers white or pinkish, 6-7 mm. long; corolla urn-shaped, the 5 short lobes spreading; stamens 10, the anthers opening by false terminal pores, each with 2 curved, reflexed, horn-like appendages; ovary superior, pubescent.
Fruits	Fruit a reddish-brown, globose berry, 7-10 mm. in diameter.	Fruit a berry, depressed, 6-8 mm. broad.

	Arctostaphylos nevadensis	Arctostaphylos columbiana
Identification notes	The fruit is the best feature for distinguishing A. nevadensis from A. uva-ursi. The fruit of A. nevedensis is reddish with splotches of brown or black; that of A. uva-ursi is bright red. Without fruit, look at the leaf color. The leaves of A. nevadensis are bright green on both surfaces; those of A. uva-ursi are dark green on the upper surface and light green on the lower.
Flowering time	May-July	May-July
Habitat	Dry forest edge and openings from middle elevations in the mountains to the subalpine.	Dry forest openings and rocky slopes, mostly at low elevations.
Distribution	Occurring on both sides of the Cascades crest in Washington; Washington to California, east to Nevada. [WildflowerSearch map] [BONAP county map]	Occurring chiefly west of the Cascades crest and in the Columbia River Gorge in Washington; British Columbia to California. [WildflowerSearch map] [BONAP county map]
Origin	Native	Native
Conservation status	Not of concern	Not of concern
Sibling taxa	A. columbiana, A. ×media, A. nevadensis × A. patula, A. patula, A. uva-ursi	A. ×media, A. nevadensis, A. nevadensis × A. patula, A. patula, A. uva-ursi
Subordinate taxa	A. nevadensis ssp. nevadensis
Web links	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Arctostaphylos columbiana

pinemat manzanita

bristly manzanita

Habit

Shrubs with spreading to decumbent stems often forming large mats or mounds, the branch tips to 2 dm. tall; bark brownish-red, stems puberulent, sometimes glandular.

Erect or spreading shrubs 1-3 m. tall, simple at the base, the old branches with purplish-red bark; young twigs grayish-pubescent, also with scattered bristles which may be glandular.

Leaves

Leaves oblong to spatulate, the tips acute or abruptly pointed;

blades 1.5-3 cm. long, glabrous to puberulent on the margins and mid-rib, the base acute or narrowed abruptly to petioles 2-5 mm. long.

Leaves alternate, ovate or lanceolate to elliptic, the blades 2-5 cm. long and 1-2.5 cm. broad, pointed, finely grayish-puberulent, the petioles 5 cm. long.

Flowers

: Inflorescence a terminal, few-flowered raceme, each flower subtended by a bract about equal to the pedicel;

flowers pink, about 5 mm. long;

corolla urn-shaped, with 5 short lobes;

stamens 10, the anthers opening by false terminal pores, each with 2 curved, reflexed, horn-like appendages;

ovary superior, pubescent.

Inflorescence usually paniculate, densely pubescent, each flower subtended by a lanceolate bract 5-15 mm. long, exceeding the pedicels;

flowers white or pinkish, 6-7 mm. long;

corolla urn-shaped, the 5 short lobes spreading;

stamens 10, the anthers opening by false terminal pores, each with 2 curved, reflexed, horn-like appendages;

ovary superior, pubescent.

Fruits

Fruit a reddish-brown, globose berry, 7-10 mm. in diameter.

Fruit a berry, depressed, 6-8 mm. broad.

The fruit is the best feature for distinguishing A. nevadensis from A. uva-ursi. The fruit of A. nevedensis is reddish with splotches of brown or black; that of A. uva-ursi is bright red. Without fruit, look at the leaf color. The leaves of A. nevadensis are bright green on both surfaces; those of A. uva-ursi are dark green on the upper surface and light green on the lower.

Flowering time

May-July

Habitat

Dry forest edge and openings from middle elevations in the mountains to the subalpine.

Dry forest openings and rocky slopes, mostly at low elevations.

Distribution