OregonFlora: Polypodium scouleri vs. Polypodium calirhiza

	Polypodium scouleri	Polypodium calirhiza
	coast polypody, leather-leaved polypody	nested polypody
Stems	stout, white-pruinose, densely clothed in scales; scales brown, longtoothed; margins often pale; taste bland.	moderately stout, often white-pruinose, with dense brown lanceolate scales; taste slightly bitter and lacking strong licorice flavor.
Leaves	to 60 cm; petiole to 25 × 0.2 cm.	to 30 cm long; petiole slender; to 13 × 0.2 cm; smooth.
Blades	pinnatifid, narrowly ovate; stiff and leathery, dark green; to 35 × 8 cm, with relatively few pairs of robust pinnae; adaxial surface glabrous; scales on abaxial surface scattered; broad; convex, brown, with pale margins.	pinnatifid; to 18 × 8 cm, narrowly ovate to oblong, often slightly leathery; adaxial surface pubescent with whitish multicellular hairs (at least along rachis), lanceolate scales usually present on abaxial surface of rachis, usually more than 3 cells wide.
Pinnae	oblong; up to 9 × 2 cm; margin usually subentire; apex rounded to acute; some veins anastomosing, forming indistinct areolae along the pinnae.	narrow, lanceolate to oblong; to 3 × 0.6 cm; margin serrate; apex usually acute, not long-attenuate; veins anastomosing or not, forming areolae on some pinnae.
Sori	round; large; over 3 mm, often absent on proximal pinnae; sporangiasters absent.	less than 3 mm; sporangiasters absent.
2n	=74.	=148.

	Polypodium scouleri	Polypodium calirhiza
Distribution	[WildflowerSearch map] [BONAP county map]	[WildflowerSearch map] [BONAP county map]
Discussion	Epiphytic and on rocks. 0–600 m. CR, Est. WA, CA; north to British Columbia, south to northern Mexico. Native.	On cliffs and rocks/gravel near the coast. 0–200 m. Est. CA; south to northern Mexico. Native.
Source	Flora of Oregon, volume 1, page 96 Duncan Thomas	Flora of Oregon, volume 1, page 95 Duncan Thomas
Sibling taxa	P. amorphum, P. calirhiza, P. glycyrrhiza, P. hesperium	P. amorphum, P. glycyrrhiza, P. hesperium, P. scouleri
Synonyms		Polypodium californicum
Web links	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Polypodium scouleri

Polypodium calirhiza

coast polypody, leather-leaved polypody

nested polypody

Stems

stout, white-pruinose, densely clothed in scales;

scales brown, longtoothed;

margins often pale;

taste bland.

moderately stout, often white-pruinose, with dense brown lanceolate scales;

taste slightly bitter and lacking strong licorice flavor.

Leaves

to 60 cm;

petiole to 25 × 0.2 cm.

to 30 cm long;

petiole slender; to 13 × 0.2 cm; smooth.

Blades

pinnatifid, narrowly ovate; stiff and leathery, dark green; to 35 × 8 cm, with relatively few pairs of robust pinnae;

adaxial surface glabrous;

scales on abaxial surface scattered; broad; convex, brown, with pale margins.

pinnatifid; to 18 × 8 cm, narrowly ovate to oblong, often slightly leathery;

adaxial surface pubescent with whitish multicellular hairs (at least along rachis), lanceolate scales usually present on abaxial surface of rachis, usually more than 3 cells wide.

Pinnae

oblong; up to 9 × 2 cm;

margin usually subentire;

apex rounded to acute; some veins anastomosing, forming indistinct areolae along the pinnae.

narrow, lanceolate to oblong; to 3 × 0.6 cm;

margin serrate;

apex usually acute, not long-attenuate;

veins anastomosing or not, forming areolae on some pinnae.

Sori

round; large; over 3 mm, often absent on proximal pinnae;

sporangiasters absent.

less than 3 mm;