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Lomatium packardiae

broad-fruit lomatium, broad nineleaf lomatium, nineleaf lomatium, Packard's lomatium

Henderson's biscuitroot, Henderson's lomatium

Habit Plants acaulescent, subacaulescent, or caulescent. Plants acaulescent or subacaulescent.
Roots

slender, elongate taproots or irregularly thickened, elongate taproots with an elongate and slender upper portion 1–6 cm.

globose or irregularly thickened tubers with elongate upper section < 1 cm and basal section abruptly tapered to a slender, elongate taproot, or irregularly thickened tubers with elongate upper section < 1 cm and bifurcate basal section that is tapered along each axis to slender taproot.

Caudices

simple, 2–3-branched or multicipital;

basal leaf sheaths from previous years weathering into a sparse thatch of a few; loose fibers or chaffy scales at base of stem or pseudoscape, previous year’s peduncles sometimes persistent in thatch as gray stalks.

simple or 2–3-branched;

basal leaf sheaths from previous years weathering into a sparse thatch of a few; loose fibers or chaffy scales at base of pseudoscape.

Stems

absent or 10–20 cm; pseudoscapes absent or < 1 cm, obscured by leaf sheaths and thatch.

absent or < 3 cm; pseudoscapes 1–3 cm, subterranean, obscured by leaf sheaths.

Basal leaves

biternate or 1–3-pinnate, glabrous, glabrate or sparsely puberulent or scaberulose along midnerves and margins, sparsely scabrous or pubescent below and glabrate or pubescent on midnerves above;

primary leaflets ternate or pinnate, 0–6 secondary leaflet pairs along each rachis;

laterals half to subequal to central primary in length;

secondaries pinnate; pinnatifid, ternate or entire;

tertiaries absent; entire; pinnatifid (2–3-lobed), ultimate apical lobes linear, narrowly elliptic, or narrowly oblong, 10–120 × 0.4–12 mm;

tips acute or acuminate, mucronulate or not.

glabrous;

primary leaflets pinnate, 1–4 secondary leaflet pairs along each rachis;

laterals less than half of central primary in length;

secondaries pinnate or pinnatifid (3–6-lobed);

tertiaries absent or entire, ultimate apical lobes elliptic; oval, obovate, 2–4.5 × 1.5–3 mm;

tips obtuse, subacute, rounded, mucronulate.

Cauline leaves

0–2, similar to basal.

0–1, similar to basal.

Inflorescences

peduncles 1–6, 10–35 cm, glabrate or sparsely to moderately scaberulose or puberulent;

involucral bracts 0;

rays 3–25, 1.5–12 cm, glabrate, sparsely scaberulose, or puberulent;

involucel bractlets 0–12, linear or lanceolate, 1–6 × 0.1–0.4 mm, glabrous; entire;

margins narrowly scarious or not;

umbellets 10–70-flowered;

pedicels 2–8 mm, glabrous, glabrate, scaberulose on ridges.

peduncles 1–6, 6–13 cm, glabrous;

involucral bracts 0;

rays 2–6, 0.5–2(2.5) cm, glabrous;

involucel bractlets 3–6, linear, lanceolate, narrowly elliptic, 2–4 × 0.3–0.8 mm;

margins narrowly scarious;

umbellets 5–15-flowered;

pedicels 1–5 mm.

Flowers

petals ochroleucous or yellow;

anthers ochroleucous or yellow.

petals ochroleucous or yellow;

anthers ochroleucous or yellow.

Fruits

oblong, narrowly oblong, elliptic or obovate, 7–16 mm, glabrous, glabrate, or sparsely scaberulose or puberulent;

body 1.3–4 mm wide;

wings 0.7–3 mm; thin;

vittae 1 in intervals, 2 on commissure.

ovate; oval, or orbicular, 4–6 mm, glabrous;

body 1–2 mm wide;

wings thin, 0.2–0.8 mm;

vittae 1 in intervals, 2 on commissure.

Lomatium triternatum

Lomatium hendersonii

Distribution
[WildflowerSearch map]
[BONAP county map]
[WildflowerSearch map]
[BONAP county map]
Discussion

Diverse habitat types but most commonly on open slopes, meadows, rocky hillsides. Flowering Feb–Jul. 0–2600 m. BR, BW, Col, CR, ECas, Lava, Owy, Sisk, WV. CA, ID, NV, WA. north to British Columbia, northeast to Alberta, east to CO, southeast to AZ. Native.

Lomatium triternatum is a complex with many ecotypes and morphotypes. Numerous varieties have been named, and some have been delimited at the species level. The ecotypes that have been named include: 1) acaulescent or caulescent plants with pubescent, short narrow apical leaflets lobes (1–4 mm wide) [= L. packardiae]; 2) caulescent plants with short narrow apical leaflets and lobes (1–2 mm wide) that are variously pubescent [= L. tamanitchii; L. triternatum var. brevifolium]; 3) caulescent plants with broad, short leaflets (10–50 × 2–8 mm) [= L. anomalum]; 4) caulescent plants from the west side of the Cascades with very long and broad leaflets (50–120 × 1–12 mm) [= L. triternatum var. macrocarpum]; 5) plants with strictly ternate lateral leaflets with lobes long and narrow (20–90 × 0.4–1.5 mm) and fruit wings (2–3 mm) equal to or wider than the body [= L. simplex]. All of these forms grade into each other imperceptibly, and often more than one form can be found on a herbarium sheet. Although Lesica and Kittlelson (2013) make a robust argu­ment for the separation of L. anomalum from L. triternatum, the condition of the types of L. anomalum prevents the reliable assignment of this name to any of the named ecotypes of L. triternatum. The types are in a mature fruiting stage with dried and withered leaves. Additionally, the leaves are fragmented, and the length and width of the leaflet lobes cannot be determined. The name needs epitypification before it can be applied to forms in the remainder of the range of L. triternatum, regard­less of how other workers have applied this name to specific morphological variants in the past.

Open gravelly or rocky slopes and flats in scrubland and woodland, on igneous-derived substrates. Flowering Mar–Jun. 400–2200 m. BR, BW, ECas, Lava, Owy. CA. Native.

The type publication of Lomatium hendersonii stated that the flowers were white. Mathias did not see the types and retained Coulter and Rose’s interpretation. The types of Leptotaenia leibergii (basionym of L. roseanum) were reported to be yellow-flowered. If it can be confirmed that the types of L. hendersonii were white and that there are extant populations in the John Day River Valley that are white-flowered L. hendersonii, the status of this taxon would need to be re-evaluated. Lomatium hendersonii is found mostly in the mountains of central Oregon, east of the Cascades and southwest of the Wallowas with disjunct outliers in southern Oregon and northern California. The reports from Owyhee County, Idaho, and Washoe County, Nevada, are likely misidentifications of L. roseanum. Lomatium hendersonii is so rarely collected that the characters of the mature fruit are poorly known. More specimens of this taxon in fruiting stage are needed to determine if the wing and body are narrower than those in L. roseanum.

Source Flora of Oregon, volume 2, page 126
Jason Alexander
Flora of Oregon, volume 2, page 118
Jason Alexander
Sibling taxa
L. ambiguum, L. bentonitum, L. bradshawii, L. brunsfeldianum, L. californicum, L. canbyi, L. columbianum, L. cookii, L. cous, L. cusickii, L. dissectum, L. donnellii, L. engelmannii, L. erythrocarpum, L. farinosum, L. foeniculaceum, L. gormanii, L. greenmanii, L. hallii, L. hendersonii, L. howellii, L. klickitatense, L. laevigatum, L. leptocarpum, L. macrocarpum, L. martindalei, L. minus, L. nevadense, L. nudicaule, L. ochocense, L. oreganum, L. papilioniferum, L. pastorale, L. peckianum, L. piperi, L. ravenii, L. rollinsii, L. roseanum, L. serpentinum, L. suksdorfii, L. triternatum, L. utriculatum, L. vaginatum, L. watsonii
L. ambiguum, L. bentonitum, L. bradshawii, L. brunsfeldianum, L. californicum, L. canbyi, L. columbianum, L. cookii, L. cous, L. cusickii, L. dissectum, L. donnellii, L. engelmannii, L. erythrocarpum, L. farinosum, L. foeniculaceum, L. gormanii, L. greenmanii, L. hallii, L. howellii, L. klickitatense, L. laevigatum, L. leptocarpum, L. macrocarpum, L. martindalei, L. minus, L. nevadense, L. nudicaule, L. ochocense, L. oreganum, L. papilioniferum, L. pastorale, L. peckianum, L. piperi, L. ravenii, L. rollinsii, L. roseanum, L. serpentinum, L. suksdorfii, L. triternatum, L. utriculatum, L. vaginatum, L. watsonii
Synonyms Lomatium alatum, Lomatium anomalum, Lomatium brevifolium, Lomatium packardiae, Lomatium platycarpum, Lomatium robustius, Lomatium simplex, Lomatium simplex var. leptophyllum, Lomatium simplex var. simplex, Lomatium triternatum ssp. platycarpum, Lomatium triternatum ssp. triternatum, Lomatium triternatum var. alatum, Lomatium triternatum var. anomalum, Lomatium triternatum var. brevifolium, Lomatium triternatum var. macrocarpum, Lomatium triternatum var. triternatum
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