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Bolander's spikerush

sedge family

Habit Plants perennial, densely cespitose; rhizomes caudex-like, mostly hidden by culms and roots, short, 1.5–3 mm thick, internodes very short. Herbs annual or perennial; cespitose or not, rhizomatous or not, rarely stoloniferous.
Culms

subterete, 10–30 cm × 0.3–0.5 mm.

usually triangular, occasionally terete, rarely compressed or quadrangular; solid or pithy.

Leaves

basal and/or cauline, alternate, usually 3-ranked, bases forming cylindric sheaths enclosing stem;

margins fused;

ligules often present;

blades frequently absent from some basal leaves, rarely from cauline leaves; when present flat, folded, rolled, or terete, linear.

Spikes

ovoid, 3–8 × 2–3 mm;

proximal scale clasping; entire, subproximal scale with flower; floral scales 8–30, 4–5 per mm of rachilla, 2–3 × 1.5 mm;

apex entire; acute, often keeled in distal part of spike.

with a shortened axis; floral scales 1–many, crowded, spirally arranged, sometimes 2-ranked, usually each subtending a single flower or perigynium, sometimes proximal and/or distal scales empty; perigynia if present closed or open down one side, subtending and enclosing a very short rachilla-bearing 1-pistillate flower (Carex), sometimes also (0)3 staminate flowers and empty scales (Kobresia);

spikes aggregated into secondary inflorescences that are panicles, often modified and sometimes consisting of a single spike; secondary inflorescences usually subtended by a foliaceous or scale-like or bristle-like bract.

Flowers

perianth bristles 3–6, often unequal; from rudimentary to 50% of achene length;

anthers 0.9–1.4 mm;

stigmas 3.

bisexual (unisexual in Carex and Kobresia);

perianth absent or consisting of (1)3–6(30) bristles and/or scales;

stamens usually (1)3, rarely more;

ovary with 1–3(4) carpels, fused; locule 1;

style undivided or branched;

stigmas 2–3(4).

Fruits

achenes, usually trigonous or biconvex.

Achenes

slightly to greatly compressed-trigonous, rarely thickly lenticular, 0.9–1.2 × 0.65–0.8 mm.

Distal leaf sheaths

persistent, not splitting;

apex obtuse, rarely hardened, lacking a tooth.

Tubercles

flat to shallowly pyramidal; lower than wide, 0.1–0.3 × 0.4–0.65 mm.

Eleocharis bolanderi

Cyperaceae

Distribution
[WildflowerSearch map]
[BONAP county map]
[BONAP county map]
Discussion

Shallow, rocky, ephemeral streams. 1100–2100 m. BR, BW, Owy. CA, NV, ID; east to CO. Native.

Eleocharis bolanderi is easily recognized in summer when it forms masses of dry, brown, dormant and apparently dead culms in dry stream beds. It is easily pulled up to expose its cespitose habit. The virtually flat tubercles are particularly distinctive.

Cosmopolitan. Approximately 100 genera; 16 genera treated in Flora.

Cyperaceae (except Eleocharis) is characterized by unusual holocentric chromosomes, in which spindle fibers attach throughout the length of the chromosome, not at discrete centromeres. Also, reduction of chromosome number occurs at the second division of meiosis. These traits are shared with the related family Juncaceae but not by other vascular plants. Chromosome numbers in sedges are often very high, but it is unclear if this represents polyploidy or fragmentation of holocentric chromosomes. When collecting sedges, get the rhizomes (if present) because some species are distinguished by their cespitose or rhizomatous growth form. If possible, collect extra inflorescences because checking floral traits can be destructive. In some genera, species identification requires mature fruits.

Source Flora of Oregon, volume 1, page 240
Barbara Wilson, Richard Brainerd, Nick Otting
Flora of Oregon, volume 1, page 167
Sibling taxa
E. acicularis, E. bella, E. decumbens, E. engelmannii, E. erythropoda, E. macrostachya, E. obtusa, E. ovata, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. rostellata, E. suksdorfiana
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