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California wood fern, coastal wood fern, marginal wood fern

Stems

short-creeping or erect.

Leaves

up to 90 × 30 cm, monomorphic, remaining green throughout winter;

petioles 25–33% of leaf length, with scattered light brown scales.

Blades

ovate-lanceolate, pinnate-pinnatifid to 2-pinnate at leaf base, with conspicuous glands.

Pinnae

deltate, with basal basiscopic and basal acroscopic pinnules approximately equal in length; pinnule margins serrate and may be bristle-tipped.

Sori

round; on ultimate segments midway between midvein and margin;

indusia reniform;

glands present when young and fading with maturity; spores brown.

Dryopteris celsa

Dryopteris arguta

Distribution
[WildflowerSearch map]
[BONAP county map]
[WildflowerSearch map]
[BONAP county map]
Discussion

Open woodlands and shaded slopes. 0–1600 m. Casc, CR, Est, Sisk, WV. CA, WA; north to British Columbia, southeast to AZ. Native.

Dryopteris arguta occurs in several variable, unnamed morphological forms. Only the most common of these is found in Oregon, and it is described here. Dryopteris arguta is somewhat similar morphologically to the eastern species D. marginalis, to which it may be closely related. Unlike most North American Dryopteris, it is not known to form any hybrids with other species.

Source Flora of Oregon, volume 1, page 73
Emily Sessa
Sibling taxa
D. arguta, D. expansa, D. filix-mas
D. expansa, D. filix-mas
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