Cotula coronopifolia |
Cotula |
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brass buttons |
cotula, waterbuttons |
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Habit | Plants perennial, 5–45 cm; usually stoloniferous. | Herbs annual or perennial; fibrous-rooted or stoloniferous. |
Stems | decumbent to erect; simple to well branched, glabrous or minutely glandular. |
1–several, decumbent to erect or spreading; simple or branched, glabrous to villous or minutely glandular. |
Leaves | linear to linear-lanceolate, 1–7 cm, bases auriculate and forming a membranous sheath around stem; margins entire or few-lobed; lobes linear; tips acute; surfaces glabrous, sometimes gland-dotted, sessile. |
principally cauline, alternate; simple to pinnately toothed or lobed, or pinnately compound; surfaces glabrous to villous or minutely gland-dotted. |
Inflorescences | solitary terminal heads. |
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Peduncles | 1–9 cm. |
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Involucres | hemispheric, 3–4 mm. |
spreading in fruit. |
Receptacles | flat to convex; paleae 0. |
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Ray florets | 0, or rarely 1–3 series of 8–80+ pistillate florets with corollas 0. |
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Disc florets | corollas 1–1.5 mm, equaling or exceeding involucres, yellow. |
corollas tubular, slightly widened distally; lobes 4–5, triangular; erect; stamens 4–5, not exserted; styles ? 1 mm. |
Phyllaries | 3–5-veined; surfaces glabrous; inner obtuse; outer obtuse or acute. |
in 2–3 equal series; lance-elliptic; margins membranous, 1–5-veined, glabrous or villous; inner obtuse; outer obtuse or acute. |
Fruits | 0.7–1.8 mm, glabrous or minutely glandular, brown, winged or not. |
obovoid or oblong, compressed, with 2 marginal veins, sometimes wing-margined; pappi 0. |
Heads | disciform, rarely radiate or discoid. |
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2n | =20. |
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Cotula coronopifolia |
Cotula |
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Distribution | ||
Discussion | Beaches, tide marshes, moist fields, disturbed areas. Flowering Mar–Dec. 0–200 m. Est. CA, NV, WA; north to British Columbia, southeast to AZ, south to Mexico; Africa, Australia, Europe, South America. Exotic. |
Africa, Australia, Europe, North America, Pacific Islands, South America. 55 species; 2 species treated in Flora. A single collection of Cotula mexicana has been made in Oregon, in 2009, from closely mowed greens of a golf course in Morrow County, where it was probably introduced in turf from California. It has a matted growth form, pinnately lobed leaves, and floral peduncles about as long as the leaves. Thus far it appears to be limited to this specialized type of cultivated habitat. |
Source | Flora of Oregon, volume 2, page 234 Kenton Chambers |
Flora of Oregon, volume 1 |
Sibling taxa | ||
Subordinate taxa | ||
Web links |
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