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Nuttall's toothwort, slender toothwort

Holmgren's bittercress

Habit Plants perennial, glabrous or pubescent; rhizomes slender, tuberous, segmented, producing fragile segments, ovoid, oblong, or cylindrical, 2–5 mm in diameter, nodes fleshy. Plants perennial, glabrous; rhizomes slender, 0.5–1 mm in diameter.
Stems

erect, 0.5–2(3) dm; simple, bases above ground much wider than attachment to rhizome.

erect to ascending, 15–20 cm, bases above ground as wide as attachment to rhizomes.

Rhizomal leaves

simple or 3(5)-foliolate; (3)4–20(25) cm;

blades or terminal leaflets reniform, suborbicular; ovate, or oblong; (0.9)1.3–4(5.2) × (0.8)1.2–5(7) cm, often with purple spots, bases cordate to obtuse;

margins apiculate-dentate or 5–7-lobed.

absent.

Cauline leaves

1–3, pinnately or palmately compound, 3(5)-foliolate, bases not auriculate;

terminal leaflets ovate, oblong, or linear; (0.5)1–3.5(6) cm;

lateral leaflets smaller; entire, dentate, or lobed.

3 or 4, pinnately or palmately compound, 3- or 5-foliolate, bases not auriculate;

terminal leaflets obovate to lanceolate or linear in uppermost leaves, 1.3–2.3 × 0.2–0.8 cm;

lateral leaflets linear to linear-lanceolate, 0.9–1.5 × 0.1–0.3 cm;

margins entire.

Inflorescences

bracts 0, fruiting pedicels ascending to divaricate, 1–3.5 cm.

bracts 0, fruiting pedicels ascending to suberect, 5–14 mm.

Flowers

sepals erect, oblong, 3.5–5 mm;

lateral pair saccate;

petals obovate, 10–15 × 4–7.5 mm, purple to pale pink, rarely white, not clawed;

tips rounded;

ovules 8–16 per ovary;

styles 4–8 mm.

sepals erect, oblong, 1.2–1.5 mm;

lateral pair not saccate;

petals oblanceolate, 2–2.5 × 0.8–1 mm, white;

tips rounded, not clawed;

ovules 16–24 per ovary;

styles 0.5–0.7 mm.

Fruits

25–56 × 2–2.3 mm, glabrous.

10–20 × ~1 mm, glabrous.

Seeds

oblong, 2–2.5 × 1.4–1.6 mm.

oblong, 1.2–1.5 × 0.8–1 mm, brown.

Cardamine nuttallii

Cardamine holmgrenii

Distribution
[WildflowerSearch map]
[BONAP county map]
[BONAP county map]
Discussion

Damp woods, shaded bottomlands and hillsides, mossy slopes, streamsides. Flowering Mar–Jul. 0–2000 m. Casc, Col, CR, ECas, Est, Sisk, WV. CA, WA; north to British Columbia. Native.

Boggy slopes. Flowering Jul–Aug. 2000–2200 m. BW. Native. Endemic to Oregon.

This species is known only from the type locality in the Blue Mountains of Baker County.

Source Flora of Oregon, volume 2 draft
Ihsan Al-Shehbaz
Flora of Oregon, volume 2, page 465
Ihsan Al-Shehbaz
Sibling taxa
C. angulata, C. bellidifolia, C. breweri, C. californica, C. cordifolia, C. flexuosa, C. hirsuta, C. holmgrenii, C. occidentalis, C. oligosperma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica
C. angulata, C. bellidifolia, C. breweri, C. californica, C. cordifolia, C. flexuosa, C. hirsuta, C. nuttallii, C. occidentalis, C. oligosperma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica
Synonyms Cardamine nuttallii var. covilleana, Cardamine nuttallii var. dissecta, Cardamine nuttallii var. gemmata, Cardamine nuttallii var. nuttallii, Cardamine pulcherrima, Cardamine pulcherrima var. pulcherrima, Cardamine pulcherrima var. tenella, Dentaria gemmata, Dentaria tenella, Dentaria tenella var. pulcherrima, Dentaria tenella var. quercetorum, Dentaria tenella var. tenella
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