Cardamine nuttallii |
Cardamine holmgrenii |
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Nuttall's toothwort, slender toothwort |
Holmgren's bittercress |
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Habit | Plants perennial, glabrous or pubescent; rhizomes slender, tuberous, segmented, producing fragile segments, ovoid, oblong, or cylindrical, 2–5 mm in diameter, nodes fleshy. | Plants perennial, glabrous; rhizomes slender, 0.5–1 mm in diameter. |
Stems | erect, 0.5–2(3) dm; simple, bases above ground much wider than attachment to rhizome. |
erect to ascending, 15–20 cm, bases above ground as wide as attachment to rhizomes. |
Rhizomal leaves | simple or 3(5)-foliolate; (3)4–20(25) cm; blades or terminal leaflets reniform, suborbicular; ovate, or oblong; (0.9)1.3–4(5.2) × (0.8)1.2–5(7) cm, often with purple spots, bases cordate to obtuse; margins apiculate-dentate or 5–7-lobed. |
absent. |
Cauline leaves | 1–3, pinnately or palmately compound, 3(5)-foliolate, bases not auriculate; terminal leaflets ovate, oblong, or linear; (0.5)1–3.5(6) cm; lateral leaflets smaller; entire, dentate, or lobed. |
3 or 4, pinnately or palmately compound, 3- or 5-foliolate, bases not auriculate; terminal leaflets obovate to lanceolate or linear in uppermost leaves, 1.3–2.3 × 0.2–0.8 cm; lateral leaflets linear to linear-lanceolate, 0.9–1.5 × 0.1–0.3 cm; margins entire. |
Inflorescences | bracts 0, fruiting pedicels ascending to divaricate, 1–3.5 cm. |
bracts 0, fruiting pedicels ascending to suberect, 5–14 mm. |
Flowers | sepals erect, oblong, 3.5–5 mm; lateral pair saccate; petals obovate, 10–15 × 4–7.5 mm, purple to pale pink, rarely white, not clawed; tips rounded; ovules 8–16 per ovary; styles 4–8 mm. |
sepals erect, oblong, 1.2–1.5 mm; lateral pair not saccate; petals oblanceolate, 2–2.5 × 0.8–1 mm, white; tips rounded, not clawed; ovules 16–24 per ovary; styles 0.5–0.7 mm. |
Fruits | 25–56 × 2–2.3 mm, glabrous. |
10–20 × ~1 mm, glabrous. |
Seeds | oblong, 2–2.5 × 1.4–1.6 mm. |
oblong, 1.2–1.5 × 0.8–1 mm, brown. |
Cardamine nuttallii |
Cardamine holmgrenii |
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Distribution | ||
Discussion | Damp woods, shaded bottomlands and hillsides, mossy slopes, streamsides. Flowering Mar–Jul. 0–2000 m. Casc, Col, CR, ECas, Est, Sisk, WV. CA, WA; north to British Columbia. Native. |
Boggy slopes. Flowering Jul–Aug. 2000–2200 m. BW. Native. Endemic to Oregon. This species is known only from the type locality in the Blue Mountains of Baker County. |
Source | Flora of Oregon, volume 2 draft Ihsan Al-Shehbaz |
Flora of Oregon, volume 2, page 465 Ihsan Al-Shehbaz |
Sibling taxa | ||
Synonyms | Cardamine nuttallii var. covilleana, Cardamine nuttallii var. dissecta, Cardamine nuttallii var. gemmata, Cardamine nuttallii var. nuttallii, Cardamine pulcherrima, Cardamine pulcherrima var. pulcherrima, Cardamine pulcherrima var. tenella, Dentaria gemmata, Dentaria tenella, Dentaria tenella var. pulcherrima, Dentaria tenella var. quercetorum, Dentaria tenella var. tenella | |
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